Representative Publications

Leroux, M. & Townsend, S.W. (in press). Call combinations in great apes and the evolution of syntax.

Engesser, S., Holub, J.L., O'Neill, L.G., Russell, A.F. & Townsend, S.W. (2019) Chestnut-crowned babbler calls are composed of meaningless shared building blocks. Proceedings of the National Academy of Sciences of the USA 119(39)19579-19584.

Townsend, S.W., Engesser, S., Stoll, S., Zuberbühler, K. & Bickel, B. (2018). Compositionality in animals and humans. PLoS Biology 16(8):e2006425.

Collier, K, Townsend, S.W. & Manser M.B. Call concatenation in wild meerkats (2017). Animal Behaviour

Engesser, S., Ridley, A. & Townsend, S.W. (2016). Meaningful call combinations and compositional processing in the Southern Pied Babbler. PNAS, 113, 5976-5981. 

Engesser, S., Crane, M.S., Savage, J.L., Russell, A.F. & Townsend, S.W. (2015). Experimental evidence for phoneme structure in the vocal system of a non-human. PLoS Biology 13(6): e1002171. 

Borgeaud, C., Alvino, M., van Leeuwen, K., Townsend, S.W. & Bshary, R. (2015). Age and sex differences in third-party rank relationship knowledge in wild vervet monkeys (Chlorocebus aethiops pygerythrus). Animal Behaviour 102, 277-284. 

Watson, S., Townsend, S.W., Schel, A.M., Wilke, C., Wallace, E. Cheng, L., West, V. & Slocombe, K.E. (2015). Vocal Learning in the Functionally Referential Food Grunts of Chimpanzees. Current Biology 25, 1-5. 

Townsend, S.W., Charlton, B.D. & Manser, M.B. (2014). Formant cues to identity and predator context in the barks of meerkats. Animal Behaviour 95, 143-149. 

Schel, A. M., Machanda, Z., Townsend, S.W., Zuberbuhler, K. & Slocombe, K.E. (2013). Chimpanzee food calls are directed at specific individuals. Animal Behaviour 85, 955-966. 

Mazzini, F.*, Townsend, S.W.*, Viranye, Z. & Range, F. (2013). Wolf howling is mediated by relationship quality rather than underlying emotional stress. Current Biology 23, 1677–1680.